Muscle Length-tension Relationship
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Muscle Fatigue & Blood Supply
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Greatest tension is developed at point B (slightly
stretched) with less tension developed at points A
(contracted) and C (stretched). |
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- Muscle Fatigue (a) muscle with intact circulation
(b) isolate muscle. Note sustained muscular contraction
may occlude local vasculature momentarily impeding blood flow
to activated muscle. Also see Pump
and Burn.
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Stretch-shortening Cycle
Muscular forces can be enhanced when the muscle is stretched
(causing eccentric tension) immediately before concentric contraction.
This stretch-shorten cycle (SSC) occurs naturally in running,
jumping, and all other activities in which muscles are suddenly
stretched by impact or other external forces. The stretch-shortening
cycle saves energy by temporarily storing potential energy through
elastic recoil resulting from an external quick stretching force.
- Plyometric vertical jumps folowing quick landings from a
0.4 m height have been found to be higher than made from a semisquat,
with or without a dip or countermovement (Asmussen & Bonde-Peterson
1976). Also see Depth
Jump Heights and Speed
of Contraction.
- Recoils or windups immediately before throwing, stiking,
or kicking can augment the muscle contraction forces to come.
- In running, the shock abosorption occuring at foot plant
contributes to greater force production at push-off (Cavagna,
Dusman & Margaria 1968).
- In sprint running, the quick stretch (0.15 to 0.1 seconds)
applied at ground contact during push-off increased force production
(Kreighbaum 1996).
- In weight training, even a one second delay at the bottom
of a bench press would result in a 55% loss of force.
Slow Oxidative (SO) fibers have
the greater elastic properties than Fast
Glycolytic (FG) (Kimi & Bosco 1978, Milner 1988)
Musculotendionous Receptors
Muscle Spindles
Muscle spindles are a type of receptor located thoughout the
muscle and situated between and parallel to individual muscle
fibers. In this position, they are stretched along with adjacent
muscle fibers causing a reflex contraction of their associated
host muscle, also known as myotatic or stretch reflex.
The knee-jerk reflex is an example. Both the stretch reflex and
recoil of stored elastice energy
used in striking or throwing movements. Muscle spindles also
are involved in motor control, providing constant monitoring
and regulation of sensorimotor function, enabling appropriate
body movement, both reflective and voluntary (Schmidt 1988, Gregor
1989).
Golgi Tendon Organs
Golgi Tendon Organs (GTO) are receptors located in tendon
merging with its muscle (myotendinous junction) and positioned
in line with the direction of muscle contraction. These receptors
are sensitive to both the passive and active pull of muscle tendon
(Gregor, 1989). The GTO inhibits the contraction of the associated
muscle (autogenic inhibition) and excites the antagonistic
muscle group. Also see Golgi
Tendon Organs Question/Answer.
Fibers Types
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Slow oxidative (SO) |
Fast Oxidative Glycolytic (FOG) |
Fast Glycolytic (FG) |
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Type I (red) |
Type IIa (white) |
Type IIb (white) |
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Speed of contraction |
Slow |
Fast |
Fast |
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Force of contraction |
Low |
High |
High |
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Anaerobic capacity |
Low |
Medium |
High |
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Aerobic capacity |
High |
Medium |
Low |
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Capillary density |
High |
Medium |
Low |
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Mitochondrial density |
High |
High |
Low |
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Motor neuron size |
Small |
Medium |
Large |
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Major substrate |
Triglycerides |
CP, Glycogen |
CP, Glycogen |
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Activity |
Prolonged low intense |
Prolonged high intense |
Short high intense |
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Average fiber percentage |
50% |
35% |
15% |
- Ratio of both types of muscle fiber varies in each individual
and each muscle.
- Ratio is thought to be genetically determined.
- Both types of fibers can improve their metabolic capabilities
through specific strength and endurance training.
- Sprinters and weight lifters have a large percentage of fast-twitch
fibers.
- Marathon runners generally have a higher percentage of slow
twitch fibers.
- Muscles that primarily maintain posture against gravity require
more endurance and generally have a higher percentage of slow-twitch
fibers.
- Muscles that produce powerful, rapid, explosive strength
movements tend to have a greater percentage of fast-twitch fibers.
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An artery branching into capillaries between three muscle
fibers |
Record of successive contractions of elbow flexor muscles |
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One end of muscle fiber showing attachment of tendon to sarcolemma |
Part of a muscle fiber specially prepared to bring out the numerous
nuclei |
Also see motor
unit recruitment.
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